OK, today when I left for work it was 44 degrees with high prospects for getting up in the 50's (bee flying weather). I spread flour on the landings of my three hives.
Aristaeus2:

Mellona:

Bermuda (the hive that isn't using it's landing but rather an upper opening as an entry)

When I came home the landing on Aristaeus2 was all cleaned off and so I couldn't see any bee footprints. The landing on Mellona was clean as well.


Bermuda had these prints but they look like they might represent squirrel paws rather than bee feet. So the mystery remains unsolved.

Well, I'll be home tomorrow and again it should be sunny and warm enough for them to fly.
Oh I hope they're ok!
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